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National Nematode Collection of New Zealand (NNCNZ)

The National Nematode Collection of New Zealand (NNCNZ) is a collection of nematode (roundworm) type specimen from New Zealand and worldwide. 

The NNCNZ at present contains 178 holotypes of New Zealand species and 19 holotypes of species from elsewhere. Samples are generally used by researchers for nematode taxonomic studies and identification. 

The National Nematode Collection of New Zealand (NNCNZ) was established in 1971 at the Entomology Division of the Department of Scientific and Industrial Research, Nelson. It was moved to Auckland when the Entomology Division was transferred there in September 1973. In 1992 the collection became a component of the New Zealand Arthropod Collection (NZAC), in the Crown Research Institute Landcare Research New Zealand Ltd, as part of a restructuring of research and amalgamation of various taxonomic activities. 

Earlier lists of specimens in the NNCNZ were published in 1972, 1973 and 1996 (Wouts & Yeates 1972; Wouts 1973a, Wouts 1996).  In the 1973 paper an historical perspective was presented. At that time the collection contained all but two of the 82 holotypes ever assigned to nematodes from New Zealand.  The holotype of Iotonchus basidontus Clark, 1960 could not be located, and the holotype of Radopholus nativus Sher, 1968 was in the nematode collection of the University of California, Riverside, California.

By 1996 Tylencholaimus chathami Yeates, 1979 had been lost in transit (G.W. Yeates, pers. comm.) and the holotype of Aranimermis giganteus Poinar & Early, 1990 had been deposited in the National Museum of New Zealand, Wellington.

Subsequently five newly described species from New Zealand had their holotypes deposited in the nematode collection of the University of California: Antarctylus humus Sher, 1973 at Riverside, and Nothocriconema pasticum Raski & Pinochet, 1976, N. spinicaudatum Raski & Pinochet, 1976, Blandicephalonema serratum Mehta & Raski, 1971, and B. pilatum Mehta & Raski, 1971 at Davis.

Since 1996 the slides of Tetleyus clarki Egunjobi, 1968b, Tylenchus areolatus Egunjobi, 1968b and T. neozealandicus Egunjobi, 1968b have become irreparably damaged and no specimens can now be found on them. Paratypes of Aranimermis giganteus Poinar & Early, 1990 are present in the NNCNZ but are not listed below because they are in vials and not on slides like the other material.

In the 40 years of its existence the Collection has steadily grown through contributions by W.C. Clark, G.W. Yeates and W.M. Wouts. Minor contributions were also made by, among others, D. Sturhan and students of W.C. Clark, D.A. Wharton and myself.  With the retirement of G.W. Yeates in 2009, the type material of species described by him, deposited at the Nematology Department, Rothamsted Experimental Station, Harpenden, Herts, England (Wouts & Yeates 1972), has now been accommodated in the New Zealand collection.

The NNCNZ at present contains 178 holotypes of New Zealand species and 19 holotypes of species from elsewhere. Each nematode species held in the NNCNZ on 31 May 2010 is listed, with the author, sex of the holotype, amount of paratype material, and NNCNZ numbers.

List of type material in the NNCNZ

Names as in the original publication. Column code: a, sex of holotype (f female, m male); b, female paratypes (n); c, male paratypes (n); d, juvenile paratypes (n); e, NNCNZ numbers identifying the paratype slides.

a b c d e
I. New Zealand species 
Acrobeles kotingotingus Yeates, 1967e f 10 9 13 1-29
Acrobeles maeneeneus Yeates, 1967e f 8 9 19 594-629
Acrobeles taraus Yeates, 1967e f 2 9 8 1378-1395
Acrobeloides ellesmerensis Yeates, 1967e f 1   13 1270-1284
Acrobeloides syrtisus Yeates, 1967e f 2   9 1203-1213
Actinolaimus zealandicus Clark, 1963d f 40 28 10 31-58
Alaimus himatangiensis Yeates, 1967c f 5 6   1259-1269
Amphorostoma saccatum Clark, 1962 f 1 1   1374-1375
Anaplectus arenicola Killick, 1964 f 18   4 77-85
Anatonchus killicki Clark, 1963e f 7   5 86-90
Anguilluloides zondagi Dale, 1967a m 7 10   1487-1494
Antarctylus humus Sher, 1973   2   4 742
Aporcelaimellus maitai Yeates, 1967f f 8   7 91-105
Aporcelaimellus taylori Yeates, 1967f f   4 2 630-635
Aranimermis giganteus Poinar & Early, 1990   5     5 vials
Arenasoma terricola Yeates, 1967f m 6 4 9 830-848
Ascaridia zelandica Clark, 1981 m 8 2   1940-1946
Axonchium coxi Yeates, 1979 f 6   4 1911-1916
Axonchium watti Yeates, 1979 f 1 1 15 1907-1910
Beleodorus spiralis Egunjobi, 1968a f 4     1376-1377
Blatticola tuapekae Dale, 1966c m 6 2   1503-1508
Campbellenchus filicauda Wouts, 1978 f 24 19 14 1826-1835
Campbellenchus poae Wouts, 1978 f 282 195 11 1792-1825
Cephalobellus costelytrae Dale, 1964b m 4 2   1434-1438
Cephalobellus fluxi Dale, 1966a m 8 3   1439-1445
Cobbonchus australis Clark, 1963e f 2 1 22 147-155
Cobbonchus chauliodus Clark, 1961b f     1 131
Cobbonchus pounamua Clark, 1961b f 2 3 12 132-146
Crassolabium australe Yeates, 1967f f 6 9 28 1141-1183
Cyathostoma cacatua Bowie, 1985a f 2 2   2012-2015
Discolaimium sabuli Yeates, 1967f f   3 7 192-201
Discolaimus arenicolus Yeates, 1967f f 3   3 652-658
Dolichodorus arenarius Clark, 1963a m 6 7 3 156-167
Dorylaimellus egmonti Yeates & Ferris, 1984 f 9   2 2016-2018
Dorylaimellus monticolus Clark, 1963d f 2     1372-1373
Dorylaimellus pastura Yeates, 1979 f 33   15 1927-1933
Dorylaimellus tahatikus Yeates, 1967f f 6 2   644-651
Dorylaimoides chathami Yeates, 1979 f 16   2 1896-1900
Doryllium australe Grandison, 1964 f 14 4 1 1184-1202
Dudekemia alpinense Bowie, 1985b f 24 7 4 2078-2090
Dudekemia hirsutus Bowie, 1985b f 16 27 2 2104-2113
Dudekemia kaorinus Bowie, 1985b m 2 7 2 2093-2102
Dudekemia prolifica Bowie, 1985b f 6 2   2066-2073
Ereptonema inflatum Yeates, 1967h f 2     667-668
Eurystomina whangae Yeates, 1967c m 6 5 6 1285-1301
Falcihasta palustris Clark, 1964b f 14 23 8 202-231
Globodera zelandica Wouts, 1984 f 72 116 44 1992-2010
Haliplectus onepui Yeates, 1967b f 6 5 14 334-358
Helicotylenchus depressus Yeates, 1967a f 6   6 245-256
Hemicycliophora chathami chathami Yeates, 1978 f 3 3 6 1884-1891
Hemicycliophora chathami major Yeates, 1978 f 20 3 8 1874-1883
Hemicycliophora halophila Yeates, 1967h f 11   2 232-244
Heterodera aucklandica Wouts & Sturhan, 1995a f 149 148 152 2174-2210
Heterodera litoralis Wouts & Sturhan, 1995b f 104 4 309 2339-2361
Heth hamatus Bowie, 1985b f 11 6   2048-2064
Iotonchus maragnus Clark, 1961a f 1 1 5 302-306
Iotonchus ophiocercus Clark, 1961a f 5 2 29 307-328
Iotonchus percivali Clark, 1961a f   1 6 329-333
Iotonchus stockdilli Yeates, 1988 f 20 9 44 2150-2173
Labronema rikia Yeates, 1967f f 2 6   636-643
Leptonchus dicephalus Yeates, 1967f f 3 7 14 168-191
Longibulbophora ammophilae Yeates, 1967d f 42   14 1057-1083
Longidorus orongorongensis Yeates, Etteger & Hooper, 1992 f 4 3 45 2123-2148
Longidorus taniwha Clark, 1963b f 2   7 359-364
Mesorhabditis littoralis Yeates, 1969 f 3 8 16 686-712
Miconchus kirikiri Yeates, 1967c f 2 3   722-726
Miconchus reflexus Yeates, 1967c f 13 3 5 59-76
Mononchoides potohikus Yeates, 1969 f 29 3 22 1084-1135
Mononchus composticola Clark, 1960 f 64 3 17 442-480
Mononchus mesadenus Clark, 1960 f 24 31 32 407-441
Mononchus propapillatus Clark, 1960 f 98 1 45 365-406
Mylonchulus arenicolus Clark, 1961c f   2   489-490
Mylonchulus psammophilus Yeates, 1967c m 17 9 8 778-807
Mylonchulus ubis Clark, 1961c f 6 4   481-488
Neoaplectana leucaniae Hoy, 1954 (lectotype) f 78 19 7 1396-1417
Oxydirus visseri Yeates, 1979 f 12 13 12 1917-1926
Pakira orae Yeates, 1967b m 15 14 22 491-541
Panagrolaimus australis Yeates, 1969 f 2 3 12 669-685
Pandurinema mowhitia Yeates, 1967b m 21 12 19 990-1035
Paratylenches halophilus Wouts, 1966 f 9 7   1418-1433
Protrellina gurri Dale, 1966c m 7 7   1478-1486
Pungentus maorium Clark, 1963d f 64   26 563-586
Radopholus cavenessi Egunjobi, 1968a f        
Radopholus nativus Sher, 1968   6     1509
Ruamowhitia orae Yeates, 1967g m 2 3 16 106-127
Scutellonema magna Yeates, 1967a f 26 13 7 1214-1258
Sectonema sica Clark, 1964a f        
Stegelleta iketaia Yeates, 1967e f 6 5 18 749-777
Stegelleta tuarua Yeates, 1967e m 1 2 3 743-748
Synonchium pacificum Yeates, 1967g f 19 42 51 849-960
Takakia waipukea Yeates, 1967g m 10 11 1 1350-1371
Takamangai waenga Yeates, 1967f f 3   5 659-666
Tetleyus clarki Dale, 1965 m 9 9 5 1446-1454
Tetleyus lissotetos Dale, 1966b m 4 3   1455-1662
Tetleyus miersi Dale, 1965 m 6 6 3 1463-1467
Tetleyus pericopti Dale, 1964a m 14 3   1468-1477
Thelastoma moko Bowie, 1985b f 11 8 2 2019-2034
Thelastoma rigo Bowie, 1985b f 10 1   2035-2045
Torumanawa wahapuensis Yeates, 1967f f 9 10 3 808-829
Trichodorus clarki Yeates, 1967d m 5 12 12 961-989
Trichodorus cottieri Clark, 1963c m 19 13 6 542-562
Trischistoma stramenti Yeates, 1972 f 3   4 1511-1517
Trissonchulus littoralis Yeates, 1967c m 13 13 22 1302-1349
Trissonchulus quinquepapillatus Yeates, 1967c m 1 8   713-721
Tylencholaimus australis Yeates, 1979 f 2     1905-1906
Tylencholaimus chathami Yeates, 1979   7     1894-1895
Tylencholaimus micronanus Yeates, 1979 f 3   1 1904
Tylencholaimus rossi Yeates, 1979 f 4   2 1901-1903
Tylenchorhynchus antarcticus Wouts & Sher, 1981 f 30 14 35 1852-1868
Tylenchorhynchus tenuicaudatus Wouts & Sher, 1981 f 64 63 51 1836-1871
Tylenchus areolatus Egunjobi, 1968b f 4     128-129
Tylenchus clarki Egunjobi, 1968a f 1     591
Tylenchus neozealandicus Egunjobi, 1968b f 4     1136-1140
Tylenchus ruatus Egunjobi, 1968b f 2     130
Tylenchus whitus Egunjobi, 1968b f 4     587-590
Wetanema hula Dale, 1967b m 6 5   1495-1502
Zelandodera coxi Wouts, 1973 f 49 26 92 1605-1649
Zelandodera nothophagi Wouts, 1973 f 108 14 65 1569-1604
Zelandodera podocarpi Wouts, 1973 f 71 10 80 1538-1568
Zeldia punua Yeates, 1967e f 2 13   727-741
II. Non-New Zealand species 
Actus neocaledonensis Yeates, 1992 f 14   31 2242-2265
Atalodera ucri Wouts & Sher, 1971   10 16 23 1040-1056
Clarkus dorsalis Yeates, 1992 f 25   40 2218-2237
Clarkus ouinnensis Yeates, 1992 f 3   1 2238-2241
Cobbonchus altitudinarum Yeates, 1992 f 7   15 2285-2301
Cobbonchus couleensis Yeates, 1992 f 5 2 7 2212-2217
Cobbonchus dzumaci Yeates, 1992 f 2   9 2302-2312
Cobbonchus orstomi Yeates, 1992 f 4   4 2313-2320
Deladenus siricidicola Bedding, 1968   7   4 1789-1791
Deladenus wilsoni Bedding, 1968   9   5 1786-1788
Discocriconemella caudaventer Orton Williams, 1979   1     1934
Eutylenchus vitiensis Orton Williams, 1979   1 1   1935
Falcihasta tropica Yeates, 1973 f 22 26 28 1707-1739
Helicobelondira hebridensis Yeates, 1973 f 4 4 2 1744-1748
Hemicycliophora vitiensis Orton Williams, 1978   2 1   1936-1938
Heterodera achilleae Golden & Klindic, 1973   10     1893
Heterodera daverti Wouts & Sturhan, 1978   32   42 1872- 1960
Heterodera solanacearum Miller & Gray, 1972   23 2 2 1518-1526
Heterodera trifolii Wouts & Sturhan, 1978 (neotype) f        
Hexactinolaimus aneityi Yeates, 1973 f 2 1   1740a-c
Hylonema ivorense Luc, Taylor & Cadet, 1978   1 2 5 1989-1991
Iotonchus candelabri Yeates, 1992 f 2 2 21 2321-2329
Iotonchus lacuplanarum Yeates, 1992 f   2 12 2330-2334
Iotonchus montanum Yeates, 1992 f 1   3 2335-2338 
Iotonchus recessus Yeates, 1992 f        
Meloidodera belli Wouts, 1973b f 8   77 1530-1537
Meloidogyne camelliae Golden, 1979   1 1 1 1941-1951 
Meloidogyne querciana Golden, 1979   1   1 1947-1948 
Mitoaxonchium basalticus Yeates, 1973 f 29   7 1772-1785 
Mononchus mauritianus Williams, 1958   1     30 
Morulaimus arenicolus Sauer, 1966   12 11   592 
Mylonchulus ananasi Yeates, 1992 f 2   3 2278-2281 
Mylonchulus ciradi Yeates, 1992 f 5   4 2276-2277 
Mylonchulus paitensis Yeates, 1992 f 4   6 2282-2284 
Mylonchulus vasis Yeates, 1992 f 15   4 2266-2275 
Neoactinolaimus proximus Yeates, 1973 f 1   2 1741-1743 
Paratylenchus gabrici Yeates, 1973 f 48 13   1749-1771 
Sarisodera africana Luc, Germani & Netscher, 1973   5 1 3 1986-1988 
Sarisodera hydrophila Wouts & Sher, 1971   29 29 15 1036-1039 
Sherodera lonicerae Wouts, 1973b   145 37 157 1651-1706 
Xiphinema monohysterum Brown, 1968   1     593 
Zeldia minor Allen & Noffsinger, 1972   1     1527 
Zeldia neoacuta Allen & Noffsinger, 1972   1     1529 
Zeldia solata Allen & Noffsinger, 1972   1     1528

References

Allen, M.V. & Noffsinger, E.M. 1972. A review of the genus Zeldia  Thorne, 1937 (Nematoda: Cephalobidae) with descriptions of seven new species.  Proceedings of the Helmithological Society of Washington 39:  206-223. 

Bedding, R.A. 1968. Deladenus wilsoni n.sp. and D. siricidicola n.sp. (Neotylenchidae), entomophagous-mycetophagous nematodes parasitic in siricid woodwasps. Nematologica 14: 515-525. 

Bowie, J.Y. 1985a. Cyathostoma cacatua n.sp. (Nematoda: Syngamidae) from a sulphur-crested cockatoo, Cacatua galerita (Aves: Psittaciformes), with a key to the species of Cyathostoma. New Zealand Journal of Zoology 12: 425-429. 

Bowie, J.Y. 1985b. New species of rhigonematid and thelastomatid nematodes from indigenous New Zealand millepedes. New Zealand Journal of Zoology 12: 485-503. 

Brown, R.H. 1968. Xiphinema monohysterum n. sp. (Nematoda: Dorylaimida) from southern New Zouth Wales. Nematologica 13: 633-637. 

Clark, W.C. 1960. New Zealand Mononchidae (Enoplida, Nematoda) I. The genera Mononchus Bastian and Prionchulus Cobb. Nematologica 5: 199-214. 

Clark, W.C. 1961a. The Mononchidae (Enoplida: Nematoda) of New Zealand. II. The Genus Iotonchus (Cobb, 1916) Altherr, 1950. Nematologica 5: 260-274. 

Clark, W.C. 1961b. The Mononchidae (Enoplida: Nematoda) of New Zealand. III. A review of the genus Cobbonchus Andrassy, 1958 with descriptions of new species. Nematologica 5: 275-284. 

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Clark, W.C. 1962. Amphorostoma saccatum n.gen. et sp. family Leptonchidae (Enoplida, Nematoda). Nematologica 7: 193-196. 

Clark, W.C. 1963a. A new species of Dolichodorus (Nematoda: Tylenchida) from coastal dune sand. New Zealand Journal of Science 6: 531-534. 

Clark, W.C. 1963b. A new species of Longidorus (Micol.) (Dorylaimida: Nematoda). New Zealand Journal of Science 6: 607-611. 

Clark, W.C. 1963c. A new species of Trichodorus (Nematoda: Enoplida) from Westland, New Zealand. New Zealand Journal of Science 6: 414-417. 

Clark, W.C. 1963d. New species of dorylaimoid nematodes belonging to the genera Pungentus Thorne and Swanger, Actinolaimus Cobb and Dorylaimellus Cobb. New Zealand Journal of Science 6: 565-576. 

Clark, W.C. 1963e. Notes on the Mononchidae (Nematoda) of the New Zealand region with descriptions of new species. New Zealand Journal of Science 6: 612-632. 

Clark, W.C. 1964a. A new species of Sectonema Thorne, 1930 (Family Nygolaimidae, Nematoda). New Zealand Journal of Science 7: 174-176. 

Clark, W.C. 1964b. Falcihasta palustris n.gen. et sp., family Belondiridae (Nematoda). New Zealand Journal of Science 7: 177-80. 

Clark, W.C. 1981. Nematode parasites of the shining cuckoo, Chrysococcyx lucidus (Avis: Cuculiformes), in New Zealand. New Zealand Journal of Zoology 8: 1-4. 

Dale, P.S. 1964a. Tetleyus pericopti n.gen. et. sp., a thelastomatid nematode from the larva of Pericoptus truncatus (Fab.) (Coleoptera: Dynastinae). New Zealand Journal of Science 7: 589-595. 

Dale, P.S. 1964b. A new species of Cephalobellus (Nematoda: Thelastomatidae) from the larva of Costelytra zealandica (Coleoptera: Melolonthinae). New Zealand Journal of Science 7: 596-601. 

Dale, P.S. 1965. Two new nematodes of the genus Tetleyus (Thelastomatidae) from insect larvae. New Zealand Journal of Science 8: 521-530. 

Dale, P.S. 1966a. Cephalobellus fluxi n.sp., a thelastomatid nematode from a tipulid larva. New Zealand Journal of Science 2: 129-134. 

Dale, P.S. 1966b. A key to nematodes of the genus Tetleyus (Thelastomatidae) and a description of Tetleyus lissotetos n.sp. New Zealand Journal of Science 9: 161-165. 

Dale, P.S. 1966c. Blatticola tuapakae and Protrellina gurri n.spp., nematode parasites of the black roach. New Zealand Journal of Science 9: 538-544. 

Dale, P.S. 1967a. Nematodes associated with the pine-bark beetle, Hylastes ater, in New Zealand. New Zealand Journal of Science 10: 222-234. 

Dale, P.S. 1967b. Wetanema hula n.gen. et sp., a nematode from the weta Hemideina thoracica (White). New Zealand Journal of Science 10: 402-406. 

Egunjobi, O.A. 1968a. Three new species of nematodes from New Zealand. New Zealand Journal of Science 11: 488-497. 

Egunjobi, O.A. 1968b. Four new species of the genus Tylenchus Bastian, 1865 (Nematoda: Tylenchida). Nematologica 13: 417-24. 

Golden, A.M. 1979. Description of Meloidogyne camelliae n.sp. and M. querciana n.sp. (Nematoda: Meloidogynidae), with SEM and host-range observations. Journal of Nematology 11: 175-189. 

Golden, A.M. & Klindic, O. 1973. Heterodera achilleae n.sp. (Nematoda: Heteroderidae) from yarrow in Yugoslavia. Journal of Nematology 5: 196-201. 

Grandison, G.S. 1964. A new species of Doryllium (Nematoda: Dorylaimida) from Auckland and Campbell Islands. New Zealand Journal of Agricultural Research 7: 169-173. 

Hoy, J.M. 1954. The biology and host range of Neoaplectana leucaniae. A new species of insect parasitic nematode. Parasitology 44: 392-399. 

Killick, J.L. 1964. A new species of the genus Anaplectus de Coninck and Schuurmans Stekhoven, 1933 (Nematoda, Plectidae). New Zealand Journal of Science 7: 165-168. 

Luc, M., Taylor, D.P. & Cadet, P. 1978. Description of a new tropical Heteroderidae, Hylonema ivorense n.gen, n.sp. and a new outlook on the family Heteroderidae (Nematoda: Tylenchida). Revue de Nématologie 1: 73-86. 

Luc, M., Germani, G. & Netscher, C. 1973. Description de Sarisodera africana n. sp. et considerations sur les relations entre les genres Sarisodera Wouts and Sher, 1971 et Heterodera A. Schmidt, 1871 (Nematoda: Tylenchida). Cahiers ORSTOM, Série Biologie 21: 35-43. 

Miller, L.I. & Gray, B.J. 1972. Heterodera solanacearum n.sp., a parasite of solanaceous plants. Nematologica 18: 404-413. 

Orton Williams, K.J. 1978. Two new species of the genus Hemicycliophora de Man, 1921 (Nematoda: Tylenchida). Revue de Nématologie 1: 197-205. 

Orton Williams, K.J. 1979. The Discocriconemella species of the Fiji Islands (Nematoda: Criconematoidea). Systematic parasitology 1: 75-82. 

Poinar, G.O. & Early, J.W. 1990. Aranimermis giganteus n.sp. (Mermithidae: Nematoda) a parasite of New Zealand mygalomorph spiders (Araneae: Arachnida). Revue de Nématologie 13: 403-410. 

Sauer, M.R. 1966. Morulaimus, a new genus of the Belonolaiminae. Nematologica 11: 609-618. 

Sher, S.A. 1968. Revision of the genus Radopholus Thorne, 1949 (Nematoda: Tylenchoidea). Proceedings of the Helminthological Society of Washington 35: 219-237. 

Sher, S.A. 1973. Antarctylus humus n.gen., n.sp. from the Subantarctic (Nematoda: Tyelenchoidea). Journal of Nematology 5: 218-221. 

Williams, J.R. 1958. Studies on the nematode soil fauna of sugar cane fields in Mauritius. I. The genus Mononchus (Trilobidae, Enoplida). Occasional Paper, Mauritius Sugar Industry Research Institute 1. 13 p. 

Wouts, W.M. 1966. Paratylenchus halophilus (Nematoda: Criconematidae) a new species from New Zealand. New Zealand Journal of Science 9: 281-286. 

Wouts, W.M. 1973a. The National Nematode Collection of New Zealand. New Zealand Journal of Science 16: 3-8. 

Wouts, W.M. 1973b. A revision of the family Heteroderidae (Nematoda: Tylenchoidea) II. The subfamily Meloidoderinae. Nematologica 19: 218-235. 

Wouts, W.M. 1978. Campbellenchinae (Nematoda: Tylodoridae), a new subfamily from Campbell Island, with a description of two new species. New Zealand Journal of Zoology 4: 213-216. 

Wouts, W.M. 1984. Globodera zelandica n.sp. (Nematoda: Heteroderidae) from New Zealand, with a key to the species of the genus Globodera. New Zealand Journal of Zoology 11: 129-135. 

Wouts, W.M. & Sher, S.A. 1971. The genera of the subfamily Heteroderinae (Nematoda: Tylenchoidea) with a description of two new genera. Journal of Nematology 1: 129-144. 

Wouts, W.M. & Sher, S.A. 1981. Two new Tylenchorhynchus species (Nematoda: Tylenchorhynchidae) from the sub-antarctic region. Nematologica 27: 253-258. 

Wouts, W.M. & Sturhan, D. 1978. The identity of Heterodera trifolii Goffart, 1932 and the description of H. daverti n.sp. (Nematoda: Tylenchida). Nematologica 24: 121-128. 

Wouts, W.M. & Sturhan, D. 1995a. Heterodera aucklandica sp.n. (Nematoda: Heteroderidae) from a New Zealand native grass, with notes on the species of the avenae group. New Zealand Journal of Zoology 22: 199-207. 

Wouts, W.M. & Sturhan, D. 1995b. Heterodera litoralis sp.n. (Nematoda: Heteroderidae) from austral glasswort, Sarcocornia quinqueflora in New Zealand. Nematologica 41 (in press). 

Wouts, W.M. & Yeates, G.W.1972. Redistribution of type material. Nematologica 18: 152-153. 

Yeates, G.W. 1967a. Studies on nematodes from dune sands. I. Tylenchida. New Zealand Journal of Science 10: 280-286. 

Yeates, G.W. 1967b. Studies on nematodes from dune sands. 2. Araeolaimida. New Zealand Journal of Science 10: 287-298. 

Yeates, G.W. 1967c. Studies on nematodes from dune sands. 3. Oncholaimidae, Ironidae, Alaimidae and Mononchidae. New Zealand Journal of Science 10: 299-321. 

Yeates, G.W. 1967d. Studies on nematodes from dune sands. 4. Diphtherophoroidea. New Zealand Journal of Science 10: 322-329. 

Yeates, G.W. 1967e. Studies on nematodes from dune sands. 5. Acrobelinae. New Zealand Journal of Science 10: 527-547. 

Yeates, G.W. 1967f. Studies on nematodes from dune sands. 6. Dorylaimoidea. New Zealand Journal of Science 10: 752-84. 

Yeates, G.W. 1967g. Studies on nematodes from dune sands. 7. Monhysterida and Chromadorida. New Zealand Journal of Science 10: 785-801. 

Yeates, G.W. 1967h. Studies on nematodes from dune sands. 8. Hemicycliophora halophila n.sp and Ereptonema inflatum n.sp. New Zealand Journal of Science 10: 802-808. 

Yeates, G.W. 1969. Three new Rhabditida (Nematoda) from New Zealand sand dunes. Nematologica 15: 115-128. 

Yeates, G.W. 1972. Trischistoma stramenti n. sp. (Nematoda: Enoplida) from leaf litter. New Zealand Journal of Science 14: 897-900. 

Yeates, G.W. 1973. Taxonomy of some soil nematodes from the New Hebrides. New Zealand Journal of Science 15: 673-697. 

Yeates, G.W. 1978. Hemicycliophora chathami n.sp. (Nematoda: Tylenchida) from Chatham Island, N.Z. with the description of two subspecies. Nematologica 24: 425-435. 

Yeates, G.W. 1979. Nine new Dorylaimida (Nematoda) from the New Zealand region. Nematologica 25: 419-438. 

Yeates, G.W. 1988. Description of Mononchoidea (Nematoda: Enoplea) in pasture soil, with description of Iotonchus stockdilli n.sp. New Zealand Journal of Zoology 14: 351-358. 

Yeates, G.W. 1992. Nematodes from New Caledonia. 1. Introduction and Mononchoidea. Fundamental and Applied Nematology 15: 101-126. 

Yeates, G.W. & Ferris, V.R. 1984. Dorylaimellus egmonti n. sp. (Nematoda: Dorylaimida) from Taranaki, New Zealand. New Zealand Journal of Zoology 11: 137-140. 

Yeates, G.W., Etteger, M. van & Hooper, D.J. 1992. Longidorus orongorongensis n.sp. (Nematoda: Dorylaimida) from subsoil of conifer/broadleaved  forest. New Zealand Journal of Zoology 19: 25-31.

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